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It is not a story mode. Its whole point is that you configure the animatronics yourself. Only a limited set of characters is available, but that does not blunt the gameplay at all.
As in the earlier games, the action takes place in a familiar location - the security office. The goal is to survive the whole night without letting anything into the room. In Custom Night you personally set how often each character shows up. That is the mode's entire hook: you can raise the difficulty until the night becomes as intense as you can handle.
Custom Night also adds an indicator the other games do not have - oxygen. It is drained by characters like the Minireenas and Yenndo. Once very little is left, the screen starts to dim. It works much like the broken ventilation effect in FNAF 3. At zero oxygen you are effectively blind: the controls still work, but you are using them by feel.
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